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Fragaria chiloensis subsp. sandwicensis

(Decne.) Staudt (1962)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Rosales Family: Rosaceae Genus: Fragaria

‘ōhelo papa [ohelo papa]

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Description

Key Characters:

Growth Form: Perennial herbs.

Stems: Producing long stolons.

Roots: Stout scaly rhizomes usually 1–4 cm long.

Leaves: Leaves compound (odd-pinnate). Leaves in a basal rosette. Blades obovate–cuneate; leaflets 2.1–5(–6) cm long, 1.2–3.5 cm wide, lateral petiolules 0.2–0.8 cm long, the terminal ones 0.2–1(–2.5) cm long. Upper surfaces glabrous or with a few scattered long hairs; lower surfaces densely silky long–strigose, somewhat coriaceous. Margins coarsely serrate, sometimes only in upper ⅔. Upper surface with conspicuously impressed veins. Petioles (3–)12–16(–21) cm long, densely long–strigose. Stipules brownish, membranous, 1–1.6 cm long, coherent to base of petioles.

Flowers: Flowers 1 to several in scapes usually ca. ½ as long as petioles at anthesis, apparently elongating after anthesis to nearly as long as petioles. Flowers bisexual (perfect) or unisexual (and then plants polygamodioecious). Pedicels usually 20–50 mm long; epicalyx lobes narrowly lanceolate, lobes alternate with the calyx lobes and persistent, (3–)5–6 mm long, densely silky long–strigose. Calyx of 5 sepals, lanceolate, distinct, imbricate, ca. 7–8 mm long, enlarging to 9–10 mm long in fruit, densely silky long–strigose. Corolla of 5 petals, white, broadly obovate, distinct, imbricate, 8–11 mm long. Stamens numerous, in 1–3 whorls, persistent; filaments slender; anthers dithecal. Ovary superior, carpels numerous, attached to the elevated conical receptacle; styles lateral.

Fruit: Fruit an aggregate consisting of numerous minute achenes borne on the surface of the enlarged fleshy receptacle; red; broadly conical; ca. 1–1.5 cm long; hirsutulous. Achenes ca. 1.5 mm long; slightly embedded in shallow pits of the receptacle. Seeds 1 per achene.

Ploidy: 2n = 56*

Habitat: Dry; disturbed areas; wet forest; and subalpine shrubland.

Elevation Range: 1,160–3,070 m.

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Lana'i Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Canad. J. Bot. 40: 883 (1962)

Other References

Wagner et al. 1990:1103 (EM, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Fragaria chiloensis subsp. sandwicensis Hualalai Preserved_Specimen Pope, W.T. Hawaii BISH 1946-05-07
2 Fragaria chiloensis subsp. sandwicensis unknown Preserved_Specimen Mann, H. 610426 BISH 1864-01-01
3 Fragaria chiloensis subsp. sandwicensis Mauna Loa ? Preserved_Specimen Rock, J.F.C. Hawaii BISH
4 Fragaria chiloensis subsp. sandwicensis East Maui, slopes of Haleakala, above Olinda Preserved_Specimen Forbes, C.N. 145 Maui BISH 1910-07-01
5 Fragaria chiloensis subsp. sandwicensis Hualalai, Kalulu Lava beds. Preserved_Specimen Rock, J.F.C. 3723 Hawaii BISH 1909-06-11
6 Fragaria chiloensis subsp. sandwicensis East Maui, Koolau Forest Reserve Shaded on ground under Vaccinium, Rubus & Metrosideros. Preserved_Specimen Harrison, B. 298 Maui BISH 1973-06-30
7 Fragaria chiloensis subsp. sandwicensis East Maui, NW of Kuiki, within Haleakala Among grass & bushes. Preserved_Specimen Degener, O. 12516 Maui BISH 1939-08-09
8 Fragaria chiloensis subsp. sandwicensis Olinda, above Sheltered grassy slope with Ranunculus. Preserved_Specimen Degener, O. 2246 Maui BISH 1927-06-15
9 Fragaria chiloensis subsp. sandwicensis Hualalai Lava fields. Preserved_Specimen Rock, J.F.C. 3752 Hawaii BISH 1909-06-11
10 Fragaria chiloensis subsp. sandwicensis Paliku, Haleakala Shrubby slope. Preserved_Specimen Degener, O. 12536 Maui BISH 1939-08-05
11 Fragaria chiloensis subsp. sandwicensis Hualalai Lava fields. Preserved_Specimen Rock, J.F.C. 3753 Hawaii BISH 1909-06-11
12 Fragaria chiloensis subsp. sandwicensis Rim of Haleakala, W of Pokahu Palaha Among bushes in grass. Preserved_Specimen Degener, O. 12513 Maui BISH 1939-08-07
13 Fragaria chiloensis subsp. sandwicensis Puu Hualalai, Kaupulehu Turfy openings. Preserved_Specimen St.John, H. 11431 Hawaii BISH 1931-12-30
14 Fragaria chiloensis subsp. sandwicensis East of Maunaanu Waterhole, Kau Forest Reserve, Kau District Growing in open to closed Acacia - Metrosideros forest with Rubus, Psychotria, and mixed fern understory. Preserved_Specimen Clarke, G. 631 Hawaii BISH 1981-10-21
15 Fragaria chiloensis subsp. sandwicensis Puu Hualalai, Puuwaawaa In moist woods. Preserved_Specimen St.John, H. 11360 Hawaii BISH 1931-12-29
16 Fragaria chiloensis subsp. sandwicensis East Maui, Makawao District, Ahupuaa of Kalialinui, upper Koolau Gap Open Styphelia-Vaccinium dominated pahoehoe W of Waikau cabin. Preserved_Specimen Higashino, P.K. 770 Maui BISH 1975-06-01
17 Fragaria chiloensis subsp. sandwicensis Haleakala Crater, Koolau gap Fog-swept grassy area. Preserved_Specimen Degener, O. 8155 Maui BISH 1927-08-11
18 Fragaria chiloensis subsp. sandwicensis Haleakala Crater, Koolau Gap E side Common at edge of thicket in meadow. Preserved_Specimen St.John, H. 21334 Maui BISH 1945-09-06
19 Fragaria chiloensis subsp. sandwicensis Makawao District, Kamaole AHU, Tr 29912/13; SW slope of Haleakala; NW of Poli Poli State Park Forest Reserve Dry exotic tree community with scattered native trees & an exotic grass understory. Preserved_Specimen Warshauer, F.R. 2665 Maui BISH 1980-06-17
20 Fragaria chiloensis subsp. sandwicensis Olinda, above Sheltered grassy slope with Ranunculus. Preserved_Specimen Degener, O. 2246 Maui BISH 1927-06-15
21 Fragaria chiloensis subsp. sandwicensis Keauhou Ranch, near Puu Kipu. Location TKY 2150 Closed Metrosideros forest with Styphelia. Preserved_Specimen Clarke, G. 80.293 Hawaii BISH 1980-02-12
22 Fragaria chiloensis subsp. sandwicensis East Maui, within Haleakala Crater Grassy dry kipuka between lava tube & spring. Preserved_Specimen Degener, O. 2266 Maui BISH 1927-08-23
23 Fragaria chiloensis subsp. sandwicensis Near hill NE of Olinda Partly shaded grassy ravine. Preserved_Specimen Degener, O. 3451 Maui BISH 1927-07-27
24 Fragaria chiloensis subsp. sandwicensis Olinda, above Sheltered grassy slope with Ranunculus. Preserved_Specimen Degener, O. 2246 Maui BISH 1927-06-15
25 Fragaria chiloensis subsp. sandwicensis Mauka of Olinda Among grass in canyon with Argyroxiphium virescens. Preserved_Specimen Degener, O. 8156 Maui BISH 1927-07-27
26 Fragaria chiloensis subsp. sandwicensis Rim of Haleakala, W of Pokahu Palaha Among bushes in grass. Preserved_Specimen Degener, O. 12513 Maui BISH 1939-08-07
27 Fragaria chiloensis subsp. sandwicensis Kilauea Forest, Puu Kipu, Kau District Patch growing on edge of summit area, slope 20-30 degrees. growing in open ground with Carex wahuensis, beneath Coprosma, Myoporum, and Metrosideros. Preserved_Specimen Davis, J. 362 Hawaii BISH 1980-09-04
28 Fragaria chiloensis subsp. sandwicensis S slope of Haleakala, in small gulch between Kahau Crater & Nawini Growing in contact of montane rainforest of Metrosideros, Acacia, Myrsine, with dry subalpine woodland of Sophora-Myoporum. Preserved_Specimen Hatheway, W.H. 462 Maui BISH 1950-11-24
29 Fragaria chiloensis subsp. sandwicensis Puu Hualalai, Puuwaawaa In moist woods. Preserved_Specimen St.John, H. 11360 Hawaii BISH 1931-12-29
30 Fragaria chiloensis subsp. sandwicensis Haleakala, SE slope of Kuiki Common in thickets at tree line. Preserved_Specimen St.John, H. 21165 Maui BISH 1945-08-26
31 Fragaria chiloensis subsp. sandwicensis Keauhou Ranch, Puu Kipu. Location TKY 2150 Closed Metrosideros forest with Styphelia. Preserved_Specimen Clarke, G. 80.293 Hawaii BISH 1980-02-12
32 Fragaria chiloensis subsp. sandwicensis Slopes of Hualalai In shrubberies. Preserved_Specimen Skottsberg, C.J.F. 602 Hawaii BISH 1922-09-25
33 Fragaria chiloensis subsp. sandwicensis Haleakala Crater, E end; 0.5 mile S of Paliku Cabin Infrequent in shaded area in heath scrub on a'a lava with Vaccinium, Polypodium, Coprosma, Styphelia, etc. Preserved_Specimen Henrickson, J.S. 3492 Maui BISH 1969-06-16
34 Fragaria chiloensis subsp. sandwicensis 50 m E of Hosmer's Grove Rocky, mesic Metrosideros-dominated narrow ravine. Preserved_Specimen Higashino, P.K. 610 Maui BISH 1975-06-12
35 Fragaria chiloensis subsp. sandwicensis Haleakala National Park, Paliku Quite frequent throughout crater in moist locations. Preserved_Specimen Olson, G.E. 46 Maui BISH 1937-08-01
36 Fragaria chiloensis subsp. sandwicensis Hualalai, Kalulu Lava beds. Preserved_Specimen Rock, J.F.C. 3724 Hawaii BISH 1909-06-11
37 Fragaria chiloensis subsp. sandwicensis Kau Forest Reserve, along trail which extends from end of access road bordering the lower boundary of Kapapala F.R Patch of about 15 plants growing along trail. Tall open Acacia koa - Metrosideros forest with Myoporum and Coprosma. Preserved_Specimen Davis, J. 550 Hawaii BISH 1981-07-02
38 Fragaria chiloensis subsp. sandwicensis Waikekeehia, foot of pali E of Haleakala Crater Turfy ledges. Preserved_Specimen St.John, H. 17743 Maui BISH 1936-12-25
39 Fragaria chiloensis subsp. sandwicensis East Maui, slopes of Haleakala Preserved_Specimen Rock, J.F.C. 8576 Maui BISH 1910-10-01
40 Fragaria chiloensis subsp. sandwicensis Kipuka Ahiu Preserved_Specimen Forbes, C.N. 902 Hawaii BISH 1915-06-21
41 Fragaria chiloensis subsp. sandwicensis Hualalai; above Kailua-Kona Preserved_Specimen Gillett, G.W. 1739 Hawaii BISH 1965-04-20
42 Fragaria chiloensis subsp. sandwicensis Kipuka (land islands in lava), near Aina Hau Preserved_Specimen Forbes, C.N. 777 Hawaii BISH 1915-06-09
43 Fragaria chiloensis subsp. sandwicensis East Maui, slopes of Haleakala Preserved_Specimen Rock, J.F.C. 8576 Maui BISH 1910-10-01
44 Fragaria chiloensis subsp. sandwicensis NW side of Koolau Gap, Haleakala Preserved_Specimen Degener, O. 12515 Maui BISH 1939-08-20
45 Fragaria chiloensis subsp. sandwicensis Slopes of Hualalai, above Puuwaawaa Preserved_Specimen Forbes, C.N. 165 Hawaii BISH 1911-06-15
46 Fragaria chiloensis subsp. sandwicensis Kipuku, near Aina Hau Preserved_Specimen Forbes, C.N. 777 Hawaii BISH 1915-06-09
47 Fragaria chiloensis subsp. sandwicensis East Maui, Haleakala Preserved_Specimen Faurie, A. 807 Maui BISH 1909-08-01
48 Fragaria chiloensis subsp. sandwicensis Paliku, Haleakala Preserved_Specimen Degener, O. 12514 Maui BISH 1939-08-05
49 Fragaria chiloensis subsp. sandwicensis Kipahulu Valley, Haleakala National Park, National Park Service Expedition III, West Camp, transect 5 Preserved_Specimen Russell, C.A. 562 Maui BISH 1984-03-03
50 Fragaria chiloensis subsp. sandwicensis Haleakala Preserved_Specimen Degener, O. 12668 Maui BISH 1927-01-01
51 Fragaria chiloensis subsp. sandwicensis Kilauea Crater Preserved_Specimen Berggren, S. Hawaii BISH 1846-11-10
52 Fragaria chiloensis subsp. sandwicensis Upper Akulele Preserved_Specimen Forbes, C.N. Lanai BISH
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